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Volgadraco

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Volgadraco
Temporal range: Late Cretaceous, Campanian
Scientific classification Edit this classification
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: †Pterosauria
Suborder: †Pterodactyloidea
Clade: †Azhdarchoidea
Family: †Azhdarchidae
Genus: †Volgadraco
Averianov et al., 2008
Type species
†Volgadraco bogolubovi
Averianov et al., 2008

Volgadraco ("Volga River dragon") is a genus of pterosaur from the Upper Cretaceous of European Russia.

Discovery and naming

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Volgadraco was named based on a lower beak (the holotype, SGU, no. 46/104a) and postcranial fragments from the early Campanian-age Rybushka Formation of Saratov, Russia. The size of this animal, and the development of blood supply in the lower jaw are intermediate between older Santonian or Turonian azhdarchids like Azhdarcho and Bakonydraco and later Maastrichtian azhdarchids like Quetzalcoatlus. Volgadraco was described in 2008 by Averianov, Arkhangelsky, and Pervushov. The type species is V. bogolubovi, the specific name honouring Russian paleontologist Nikolai Nikolaevich Bogolubov.[1]

Classification

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Averianov and colleagues initially assigned Volgadraco to the family Azhdarchidae, and considered the earlier named genus Bogolubovia to be a nomen dubium that is potentially identical to Volgadraco.[1] In contrast, Longrich and colleagues in 2018 had recovered it within the family Nyctosauridae, the cladogram of which can be seen below:[2]

Pteranodontoidea

In 2020, Averianov and Arkhangelsky reconsidered both Bogolubovia and Volgadraco as members of the family Pteranodontidae.[3] In 2022, Averianov and Kurin considered both Volgadraco and Bogolubovia as valid pteranodontids based on a new specimen of Bogolubovia.[4] In the same year, however, Volgadraco was considered to be a pteranodontian both outside the Nyctosauridae and Pteranodontidae based on phylogenetic analysis, the cladogram of which can be seen below:[5]

In 2026, Averianov and Arkhangelsky regarded the hypodigm of Volgadraco as comprising fossils of different types of pterosaurs. The holotype rostrum was retained as an azhdarchid based on the lack of curvature seen in Pteranodon, while the referred cervical vertebrae were regarded as belonging to an indeterminate Pteranodon.[6]

See also

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References

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  1. 1 2 Averianov, A.O.; Arkhangelsky, M.S.; Pervushov, E.M. (October 2008). "A New Late Cretaceous Azhdarchid (Pterosauria, Azhdarchidae) from the Volga Region". Paleontological Journal. 42 (6): 634–642. Bibcode:2008PalJ...42..634A. doi:10.1134/S0031030108060099. S2CID 129558986.
  2. ↑ Longrich, Nicholas R.; Martill, David M.; Andres, Brian; Penny, David (2018). "Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary". PLOS Biology. 16 (3) e2001663. doi:10.1371/journal.pbio.2001663. PMC 5849296. PMID 29534059.
  3. ↑ Alexander O. Averianov; Maxim S. Arkhangelsky (2020). "A large pteranodontid pterosaur from the Late Cretaceous of Eastern Europe". Geological Magazine. 158 (7): 1143–1155. doi:10.1017/S0016756820001119. S2CID 229441587.
  4. ↑ Averianov AO, Kurin AS (2022). "A new specimen of pteranodontid pterosaur Bogolubovia orientalis from the Upper Cretaceous of Penza Province, Russia". Historical Biology: An International Journal of Paleobiology. 35 (8): 1288–1296. doi:10.1080/08912963.2022.2087522. S2CID 249728681.
  5. ↑ Fernandes, Alexandra E.; Mateus, Octávio; Andres, Brian; Polcyn, Michael J.; Schulp, Anne S.; Gonçalves, António Olímpio; Jacobs, Louis L. (2022). "Pterosaurs from the Late Cretaceous of Angola". Diversity. 14 (9). 741. doi:10.3390/d14090741. hdl:10362/145845.
  6. ↑ Averianov, A. O.; Arkhangelsky, M. S. (2026). "The North American pterosaur Pteranodon in the Upper Cretaceous of Eastern Europe". Cretaceous Research 106501. doi:10.1016/j.cretres.2026.106501.