Ommatidae

Ommatidae; Temporal range: Ladinian–Recent PreꞒ Ꞓ O S D C P T J K Pg N
	Omma stanleyi
	Tetraphalerus wagneri
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Clade:	Pancrustacea
Class:	Insecta
Order:	Coleoptera
Suborder:	Archostemata
Family:	Ommatidae; Sharp & Muir, 1912

The Ommatidae are a family of beetles in the suborder Archostemata.^[1] The Ommatidae are considered the extant beetle family that has most ancestral characteristics.^[2] There are only seven extant species, confined to Australia and South America. However, the geographical distribution was much wider during the Mesozoic spanning across Eurasia, South America,^[3] and Australia, suggesting that they were widespread on Pangea.^[4] So far, over 26 extinct genera containing over 170 species of these beetles have been described.^[5] Three extant genera have been assigned to this family: Omma, Tetraphalerus and Beutelius.^[6] The family is considered to be a subfamily of Cupedidae by some authors,^[7] but have been found to be more closely related to Micromalthidae in molecular phylogenies.^[8] A close relationship with Micromalthidae is supported by several morphological characters, including those of the mandibles and male genitalia.^[9] Due to their rarity, their ecology is obscure, it is likely that their larvae feed on deadwood.^[10]

Genera

According to Kirejtshuk, 2020^[7] and subsequent literature.

Extinct genera

†Allophalerus Kirejtshuk, 2020 – Early Jurassic-Late Cretaceous (Asia)
†Asiania Lee et al., 2023^[11] – Early Cretaceous (South Korea)
†Blapsium Westwood, 1854 – Middle Jurassic (England)
†Brochocoleus Hong, 1982 – Early Jurassic-Late Cretaceous (Eurasia)
†Bukhkalius Kirejtshuk and Jarzembowski, 2020 – Burmese amber, Myanmar, mid-Cretaceous (Albian-Cenomanian)
†Burmocoleus Kirejtshuk, 2020 – Burmese amber, Myanmar, mid-Cretaceous (Albian-Cenomanian)
†Cionocoleus Ren, 1995 – Middle Jurassic-Early Cretaceous (Eurasia)
†Cionocups Kirejtshuk, 2020 – Burmese amber, Myanmar, mid-Cretaceous (Albian-Cenomanian)
†Clessidromma Jarzembowski et al., 2017 – Burmese amber, Myanmar, mid-Cretaceous (Albian-Cenomanian)
†Diluticupes Ren, 1995 – Middle Jurassic-Early Cretaceous (Eurasia)
†Echinocups Kirejtshuk and Jarzembowski, 2020 – Burmese amber, Myanmar, mid-Cretaceous (Albian-Cenomanian) (other authors consider this genus a junior synonym of Notocupes^[12])
†?Eurydictyon Ponomarenko, 1969 – Early Jurassic (Kyrgyzstan) (possibly closely related to Notocupes^[13])
†Jarzembowskiops Kirejtshuk, 2020 – Burmese amber, Myanmar, mid-Cretaceous (Albian-Cenomanian)
†Kirejtomma Li and Cai, 2021 – Burmese amber, Myanmar, mid-Cretaceous (Albian-Cenomanian)
†Lepidomma Jarzembowski et al., 2019 – Burmese amber, Myanmar, mid-Cretaceous (Albian-Cenomanian)
†Liassocupes Zeuner, 1962 – Early Jurassic (England)
†Limnomma Li and Cai in Li et al., 2021 – Middle Jurassic (China)
†Lithocupes Ponomarenko, 1966 – Late Triassic (Kyrgyzstan)
†Miniomma Li, Yamamoto and Cai, 2020 – Burmese amber, Myanmar, mid-Cretaceous (Albian-Cenomanian)
†Monticupes Ren, 1995 – Middle Jurassic-Early Cretaceous (China)
†?Notocupes Ponomarenko, 1964 – Middle Triassic-Late Cretaceous (Eurasia) (other authors have considered this genus more closely related to Cupedidae^[13])
†?Notocupoides Ponomarenko, 1966 – Late Triassic (Kyrgyzstan) (possibly closely related to Notocupes^[13])
†Odontomma Ren et al., 2006 – Early Cretaceous (China)
†Paraodontomma Yamamoto, 2017 – Burmese amber, Myanmar, mid-Cretaceous (Albian-Cenomanian)
†Pareuryomma Tan et al., 2012 – Early Cretaceous (China)
†Polyakius Kirejtshuk, 2020 – Burmese amber, Myanmar, mid-Cretaceous (Albian-Cenomanian)
†?Rhabdocupes Ponomarenko, 1966 – Late Triassic (Kyrgyzstan) (possibly closely related to Notocupes^[13])
†Rhopalomma Ashman et al., 2015 – Late Jurassic (Australia)
†Stegocoleus Jarzembowski and Wang, 2016 – Burmese amber, Myanmar, mid-Cretaceous (Albian-Cenomanian)
†Tetraphalerites Crowson, 1962 – Eocene (England)
†Zygadenia Handlirsch, 1906 – Early Jurassic-Early Cretaceous (Eurasia, ?Australia) (probably closely related to Notocupes^[13])

Extant genera

Omma Newman, 1839 – Late Triassic-Late Cretaceous (Eurasia), recent (Australia)
Tetraphalerus Waterhouse, 1901 – Middle Jurassic-Early Cretaceous (Asia), Cretaceous-recent (South America)
Beutelius Escalona et al., 2020 – Recent (Australia)

The extinct genus Fuscicupes Hong and Wang 1990 was included in the family by some authors, but was excluded by Kirejtshuk (2020).